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Steve Zissou, sea-film auteur a la Jacques Cousteau, has reason to be melancholy: his partner has been eaten, perhaps by a mythic jaguar shark, his wife may be taking up with her ex-husband, a young man appears claiming Steve is his father (Steve hates fathers), his most recent films have tanked, he's having trouble raising money for his venture to revenge his partner, and he's attracted to a pregnant reporter who prefers the pretender. At sea, in pursuit of the shark, will he escape pirates and mutiny, forge the bonds of fatherhood, place his arm around his wife, find the monster of the deep, re-establish box office hegemony, and discover a reason to smile?