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Set deep inside the icy land of Antarctica, two emperor penguins have a baby named Mumble. Hatched upon his feet, he has an amazing skill for tap-dancing. However, penguins are not supposed to tap dance. Even worse, he cannot sing. All emperor penguins are supposed to have a heart song, and he has a try at it - with awful results. Torn apart from his mom, dad, and best friend, Gloria, as a teenager who still has his adolescent feathers, his ventures off into the icy landscape. He is rescued from a leopard seal from a group of Adelie penguins, whom which the leader is Ramon. They take Mumble to their home and discover that all of their fish is being taken from an unknown 'alien' source. They ask Lovelace, an unwise and cocky leader of the Adelies, to help them. Unfortunately, he proves no help. Leading on an incredible adventure, they try to discover the threat of their food source and survival. Using courage and bravery, Mumble begins to teach everyone begins to learn that uniqueness isn't a burden, but is a gift to be treasured.